Christian Keysers - Publications#

Christian Keysers has published 104 papers. Web of science indicates: h=51, 13500 citations, Google Scholar: h=61, 27500 citations. ISI Essential Science Indicators ranks him in the top1% of neuroscientists based on citations.

In addition, Christian Keysers has published work for the general public, including blogs, chapter in trade books, but above all, a monograph 'the empathic brain', that has been published in English, German, Dutch, Turkish Japanese, Korean and Polish. This book has won the prize for best science book, by the independent publishers association, and does a notable job of explaining accurately to the general public the implications of his work.

Current Publication Summary:
Christian's most up to date publication list can be found at:
https://publons.com/researcher/2627102/christian-keysers
https://scholar.google.com/citations?hl=en&user=yogBzjEAAAAJ

Most notable peer publications:

Keysers, C., Gazzola, V., & Wagenmakers, E.-J. (2020). Using Bayes factor hypothesis testing in neuroscience to establish evidence of absence. Nature Neuroscience, 23(7), 788–799. https://doi.org/10.1038/s41593-020-0660-4

Hernandez-Lallement, J., Attah, A. T., Soyman, E., Pinhal, C. M., Gazzola, V., & Keysers, C. (2020). Harm to Others Acts as a Negative Reinforcer in Rats. Current Biology : CB, 30(6), 949-961.e7. https://doi.org/10.1016/j.cub.2020.01.017

Carrillo, M., Han, Y., Migliorati, F., Liu, M., Gazzola, V., & Keysers, C. (2019). Emotional Mirror Neurons in the Rat’s Anterior Cingulate Cortex. Current Biology, 29(8), 1301-1312.e6. https://doi.org/10.1016/j.cub.2019.03.024

Han, Y., Bruls, R., Thomas, R. M., Pentaraki, V., Jelinek, N., Heinemans, M., … Keysers, C. (2019). Bidirectional cingulate-dependent danger information transfer across rats. PLoS Biology. https://doi.org/10.1371/journal.pbio.3000524

Zaki, J., Wager, T. D., Singer, T., Keysers, C., & Gazzola, V. (2016). The Anatomy of Suffering: Understanding the Relationship between Nociceptive and Empathic Pain. Trends in Cognitive Sciences, 20(4), 249–259. https://doi.org/10.1016/j.tics.2016.02.003

Cerliani, L., Mennes, M., Thomas, R. M., Di Martino, A., Thioux, M., & Keysers, C. (2015). Increased Functional Connectivity Between Subcortical and Cortical Resting-State Networks in Autism Spectrum Disorder. JAMA Psychiatry, 72(8), 767–777. https://doi.org/10.1001/jamapsychiatry.2015.0101

Keysers, C., & Gazzola, V. (2014). Dissociating the ability and propensity for empathy. Trends in Cognitive Sciences, 18(4), 163–166. https://doi.org/10.1016/j.tics.2013.12.011

Meffert, H., Gazzola, V., den Boer, J. A., Bartels, A. A. J., & Keysers, C. (2013). Reduced spontaneous but relatively normal deliberate vicarious representations in psychopathy. Brain : A Journal of Neurology, 136(Pt 8), 2550–2562. https://doi.org/10.1093/brain/awt190

Hasson, U., Ghazanfar, A. A., Galantucci, B., Garrod, S., & Keysers, C. (2012). Brain-to-brain coupling: a mechanism for creating and sharing a social world. Trends in Cognitive Sciences, 16(2), 114–121. https://doi.org/10.1016/j.tics.2011.12.007

Keysers, C., Kaas, J. H., & Gazzola, V. (2010). Somatosensation in social perception. Nature Reviews. Neuroscience, 11(6), 417–428. https://doi.org/10.1038/nrn2833

Gazzola, V., Aziz-Zadeh, L., & Keysers, C. (2006). Empathy and the somatotopic auditory mirror system in humans. Current Biology : CB, 16(18), 1824–1829. https://doi.org/10.1016/j.cub.2006.07.072

Gallese, V., Keysers, C., & Rizzolatti, G. (2004). A unifying view of the basis of social cognition. Trends in Cognitive Sciences, 8(9), 396–403. https://doi.org/10.1016/j.tics.2004.07.002.

Keysers, C., Wicker, B., Gazzola, V., Anton, J.-L., Fogassi, L., & Gallese, V. (2004). A touching sight: SII/PV activation during the observation and experience of touch. Neuron, 42(2), 335–346. http://www.ncbi.nlm.nih.gov/pubmed/15091347.

Wicker, B., Keysers, C., Plailly, J., Royet, J. P., Gallese, V., & Rizzolatti, G. (2003). Both of us disgusted in My insula: the common neural basis of seeing and feeling disgust. Neuron, 40(3), 655–664. Retrieved from http://www.ncbi.nlm.nih.gov/pubmed/14642287.

Kohler, E., Keysers, C., Umiltà, M. A., Fogassi, L., Gallese, V., & Rizzolatti, G. (2002). Hearing sounds, understanding actions: action representation in mirror neurons. Science (New York, N.Y.), 297(5582), 846–848. https://doi.org/10.1126/science.1070311.

All publications

Han, Sichterman, Maria, Gazzola & Keysers (2020) Similar levels of emotional contagion in male and female rats. Sci. Rep. 10, 2763. https://doi.org/10.1038/s41598-020-59680-2

Gallo, Baaijen Thijs, Suttrup, Fernandes-Henriques, Paracampo, Keysers & Gazzola (2020) Bayesian statistics show a lack of change in excitability following bi-hemispheric HD-TDCS over the primary somatosensory cortices. Brain Stimulation vol. 13 640–642. https://doi.org/10.1016/j.brs.2020.02.009

Bhandari, Kirilina, Caan, Suttrup, De Sanctis, De Angelis, Keysers & Gazzola (2020) Does higher sampling rate (multiband + SENSE) improve group statistics - An example from social neuroscience block design at 3T. Neuroimage 213, https://doi.org/10.1016/j.neuroimage.2020.116731

Hernandez-Lallement, Attah, Soyman, Pinhal, Gazzola & Keysers (2020) Harm to Others Acts as a Negative Reinforcer in Rats. Curr. Biol. 30, 949-961.e7. https://doi.org/10.1016/j.cub.2020.01.017

Nostro, Ioumpa, Paracampo, Gallo, Fornari, Angelis, Gentile, Spezio, Keysers & Gazzola (2020) Neuro-computational mechanisms of action-outcome learning under moral conflict. bioRxiv 2020.06.10.143891. doi:10.1101/2020.06.10.143891. https://doi.org/10.1101/2020.06.10.143891

Keysers, Gazzola & Wagenmakers (2020) Using Bayes factor hypothesis testing in neuroscience to establish evidence of absence. Nat. Neurosci. 23, 788–799. https://doi.org/10.1038/s41593-020-0660-4

Borja Jimenez, Abdelgabar, De Angelis, McKay, Keysers & Gazzola (2020) Changes in brain activity following the voluntary control of empathy. Neuroimage 216, https://doi.org/10.1016/j.neuroimage.2020.116529

Carrillo, Han, Migliorati, Liu, Gazzola & Keysers (2019) Emotional Mirror Neurons in the Rat’s Anterior Cingulate Cortex. Curr. Biol. 29, 1301-1312.e6. https://doi.org/10.1016/j.cub.2019.03.024

Cracco, Keysers, Clauwaert & Brass (2019) Representing Multiple Observed Actions in the Motor System. Cereb. Cortex 29, 3631–3641. https://doi.org/10.1093/cercor/bhy237

Müller-Pinzler, Czekalla, Mayer, Stolz, Gazzola, Keysers, Paulus & Krach (2019) Negativity-bias in forming beliefs about own abilities. Sci. Rep. 9, 14416. https://doi.org/10.1038/s41598-019-50821-w

Kompier, Keysers, Gazzola, Lucassen & Krugers (2019) Early Life Adversity and Adult Social Behavior: Focus on Arginine Vasopressin and Oxytocin as Potential Mediators. Front. Behav. Neurosci. 13, https://doi.org/10.3389/fnbeh.2019.00143

Nastase, Gazzola, Hasson & Keysers (2019) Measuring shared responses across subjects using intersubject correlation. Soc. Cogn. Affect. Neurosci. doi:10.1093/scan/nsz037. https://doi.org/10.1093/scan/nsz037

Abdelgabar, Suttrup, Broersen, Bhandari, Picard, Keysers, De Zeeuw & Gazzola (2019) Action perception recruits the cerebellum and is impaired in patients with spinocerebellar ataxia. Brain 142, 3791–3805. https://doi.org/10.1093/brain/awz337

Han, Bruls, Thomas, Pentaraki, Jelinek, Heinemans, Bassez, Verschooren, Pruis, Lierde, Carrillo, Gazzola, Carrillo & Keysers (2019) Bidirectional cingulate-dependent danger information transfer across rats. PLoS Biol. doi:10.1371/journal.pbio.3000524. https://doi.org/10.1371/journal.pbio.3000524

Keysers, Paracampo & Gazzola (2018) What neuromodulation and lesion studies tell us about the function of the mirror neuron system and embodied cognition. Curr. Opin. Psychol. 24, 35–40. https://doi.org/10.1016/j.copsyc.2018.04.001

Amodio & Keysers (2018) Editorial overview: New advances in social neuroscience: from neural computations to social structures. Current Opinion in Psychology vol. 24 iv–vi. https://doi.org/10.1016/j.copsyc.2018.10.017

Gallo, Paracampo, Müller-Pinzler, Severo, Blömer, Fernandes-Henriques, Henschel, Lammes, Maskaljunas, Suttrup, Avenanti, Keysers & Gazzola (2018) The causal role of the somatosensory cortex in prosocial behaviour. Elife 7, https://doi.org/10.7554/eLife.32740

Thomas, De Sanctis, Gazzola & Keysers (2018) Where and how our brain represents the temporal structure of observed action. Neuroimage 183, 677–697. https://doi.org/10.1016/j.neuroimage.2018.08.056

Keysers & Gazzola (2017) A Plea for Cross-species Social Neuroscience. Curr. Top. Behav. Neurosci. 30, 179–191. https://doi.org/10.1007/7854_2016_439

Modinos, McLaughlin, Egerton, McMullen, Kumari, Barker, Keysers & Williams (2017) Corticolimbic hyper-response to emotion and glutamatergic function in people with high schizotypy: A multimodal fMRI-MRS study. Transl. Psychiatry 7, https://doi.org/10.1038/tp.2017.53

Zaki, Wager, Singer, Keysers & Gazzola (2016) The Anatomy of Suffering: Understanding the Relationship between Nociceptive and Empathic Pain. Trends Cogn. Sci. 20, 249–59. https://doi.org/10.1016/j.tics.2016.02.003

Valchev, Gazzola, Avenanti & Keysers (2016) Primary somatosensory contribution to action observation brain activity-combining fMRI and cTBS. Soc. Cogn. Affect. Neurosci. 11, nsw029. https://doi.org/10.1093/scan/nsw029

Etzel, Valchev, Gazzola & Keysers (2016) Is brain activity during action observation modulated by the perceived fairness of the actor? PLoS One 11, https://doi.org/10.1371/journal.pone.0145350

Keysers (2015) The Myth of Mirror Neurons The Real Neuroscience of Communication and Cognition. Science (80-. ). 347, 239. https://doi.org/10.1126/science.1259030

Thioux & Keysers (2015) Object visibility alters the relative contribution of ventral visual stream and mirror neuron system to goal anticipation during action observation. Neuroimage 105, 380–94. https://doi.org/10.1016/j.neuroimage.2014.10.035

Valchev, Ćurčić-Blake, Renken, Avenanti, Keysers, Gazzola & Maurits (2015) cTBS delivered to the left somatosensory cortex changes its functional connectivity during rest. Neuroimage 114, 386–397. https://doi.org/10.1016/j.neuroimage.2015.04.017

Cui, Abdelgabar, Keysers & Gazzola (2015) Responsibility modulates pain-matrix activation elicited by the expressions of others in pain. Neuroimage 114, 371–378. https://doi.org/10.1016/j.neuroimage.2015.03.034

Cerliani, Mennes, Thomas, Di Martino, Thioux & Keysers (2015) Increased Functional Connectivity Between Subcortical and Cortical Resting-State Networks in Autism Spectrum Disorder. JAMA psychiatry 72, 767–77. https://doi.org/10.1001/jamapsychiatry.2015.0101

Müller-Pinzler, Gazzola, Keysers, Sommer, Jansen, Frässle, Einhäuser, Paulus & Krach (2015) Neural pathways of embarrassment and their modulation by social anxiety. Neuroimage 119, 252–261. https://doi.org/10.1016/j.neuroimage.2015.06.036

Carrillo, Migliorati, Bruls, Han, Heinemans, Pruis, Gazzola & Keysers (2015) Repeated Witnessing of Conspecifics in Pain: Effects on Emotional Contagion. PLoS One 10, e0136979. https://doi.org/10.1371/journal.pone.0136979

Valchev, Zijdewind, Keysers, Gazzola, Avenanti & Maurits (2015) Weight dependent modulation of motor resonance induced by weight estimation during observation of partially occluded lifting actions. Neuropsychologia 66, 237–245. https://doi.org/10.1016/j.neuropsychologia.2014.11.030

Bos, Montoya, Hermans, Keysers & van Honk (2015) Oxytocin reduces neural activity in the pain circuitry when seeing pain in others. Neuroimage 113, 217–224. https://doi.org/10.1016/j.neuroimage.2015.03.049

Keysers, Meffert & Gazzola (2014) Reply: Spontaneous versus deliberate vicarious representations: different routes to empathy in psychopathy and autism. Brain 137, e273–e273. https://doi.org/10.1093/brain/awt376

Di Martino, Yan, Li, Denio, Castellanos, Alaerts, Anderson, Assaf, Bookheimer, Dapretto, Deen, Delmonte, Dinstein, Ertl-Wagner, Fair, Gallagher, Kennedy, Keown, Keysers, et al. (2014) The autism brain imaging data exchange: towards a large-scale evaluation of the intrinsic brain architecture in autism. Mol. Psychiatry 19, 659–67. https://doi.org/10.1038/mp.2013.78

Engel, Hijmans, Cerliani, Bangert, Nanetti, Keller & Keysers (2014) Inter-individual differences in audio-motor learning of piano melodies and white matter fiber tract architecture. Hum. Brain Mapp. 35, 2483–97. https://doi.org/10.1002/hbm.22343

Keysers, Perrett & Gazzola (2014) Hebbian Learning is about contingency, not contiguity, and explains the emergence of predictive mirror neurons. Behav. Brain Sci. 37, 205–206. https://doi.org/10.1017/S0140525X13002343

Keysers & Gazzola (2014) Hebbian learning and predictive mirror neurons for actions, sensations and emotions. Philos. Trans. R. Soc. B Biol. Sci. 369, 20130175–20130175. https://doi.org/10.1098/rstb.2013.0175

Cui, Arnstein, Thomas, Maurits, Keysers & Gazzola (2014) Functional magnetic resonance imaging connectivity analyses reveal efference-copy to primary somatosensory area, BA2. PLoS One 9, e84367. https://doi.org/10.1371/journal.pone.0084367

Thomas, Yatawatta & Keysers (2014). Sparse coding with a global connectivity constraint. in 2014 IEEE International Conference on Image Processing, ICIP 2014 doi:10.1109/ICIP.2014.7025830. https://doi.org/10.1109/ICIP.2014.7025830

Keysers & Gazzola (2014) Dissociating the ability and propensity for empathy. Trends Cogn. Sci. 18, 163–6. https://doi.org/10.1016/j.tics.2013.12.011

Mihov, Kendrick, Becker, Zschernack, Reich, Maier, Keysers & Hurlemann (2013) Mirroring fear in the absence of a functional amygdala. Biol. Psychiatry 73, e9-11. https://doi.org/10.1016/j.biopsych.2012.10.029

Monfardini, Gazzola, Boussaoud, Brovelli, Keysers & Wicker (2013) Vicarious Neural Processing of Outcomes during Observational Learning. PLoS One 8, e73879. https://doi.org/10.1371/journal.pone.0073879

Lee, Sun, Leung, Chu & Keysers (2013) Neural activities during affective processing in people with Alzheimer’s disease. Neurobiol. Aging 34, 706–15. https://doi.org/10.1016/j.neurobiolaging.2012.06.018

Meffert, Gazzola, den Boer, Bartels & Keysers (2013) Reduced spontaneous but relatively normal deliberate vicarious representations in psychopathy. Brain 136, 2550–62. https://doi.org/10.1093/brain/awt190

Pineda, Grichanik, Williams, Trieu, Chang & Keysers (2013) EEG sensorimotor correlates of translating sounds into actions. Front. Neurosci. 7, 203. https://doi.org/10.3389/fnins.2013.00203

Cerliani, Thomas, Jbabdi, Siero, Nanetti, Crippa, Gazzola, D’Arceuil & Keysers (2012) Probabilistic tractography recovers a rostrocaudal trajectory of connectivity variability in the human insular cortex. Hum. Brain Mapp. 33, 2005–34. https://doi.org/10.1002/hbm.21338

Hasson, Ghazanfar, Galantucci, Garrod & Keysers (2012) Brain-to-brain coupling: a mechanism for creating and sharing a social world. Trends Cogn. Sci. 16, 114–21. https://doi.org/10.1016/j.tics.2011.12.007

Gazzola, Spezio, Etzel, Castelli, Adolphs & Keysers (2012) Primary somatosensory cortex discriminates affective significance in social touch. Proc. Natl. Acad. Sci. U. S. A. 109, E1657-66. https://doi.org/10.1073/pnas.1113211109

Keysers (2012) Primate cognition: Copy that. Nature 482, https://doi.org/10.1038/482158a

Engel, Bangert, Horbank, Hijmans, Wilkens, Keller & Keysers (2012) Learning piano melodies in visuo-motor or audio-motor training conditions and the neural correlates of their cross-modal transfer. Neuroimage 63, 966–78. https://doi.org/10.1016/j.neuroimage.2012.03.038

Arnstein, Cui, Keysers, Maurits & Gazzola (2011) μ-suppression during action observation and execution correlates with BOLD in dorsal premotor, inferior parietal, and SI cortices. J. Neurosci. 31, 14243–9. https://doi.org/10.1523/JNEUROSCI.0963-11.2011

Bastiaansen, Thioux, Nanetti, van der Gaag, Ketelaars, Minderaa & Keysers (2011) Age-related increase in inferior frontal gyrus activity and social functioning in autism spectrum disorder. Biol. Psychiatry 69, 832–8. https://doi.org/10.1016/j.biopsych.2010.11.007

Atsak, Orre, Bakker, Cerliani, Roozendaal, Gazzola, Moita & Keysers (2011) Experience modulates vicarious freezing in rats: a model for empathy. PLoS One 6, e21855. https://doi.org/10.1371/journal.pone.0021855

Quadflieg, Etzel, Gazzola, Keysers, Schubert, Waiter & Macrae (2011) Puddles, parties, and professors: linking word categorization to neural patterns of visuospatial coding. J. Cogn. Neurosci. 23, 2636–49. https://doi.org/10.1162/jocn.2011.21628

Schippers & Keysers (2011) Mapping the flow of information within the putative mirror neuron system during gesture observation. Neuroimage 57, 37–44. https://doi.org/10.1016/j.neuroimage.2011.02.018

Kokal, Engel, Kirschner & Keysers (2011) Synchronized drumming enhances activity in the caudate and facilitates prosocial commitment--if the rhythm comes easily. PLoS One 6, e27272. https://doi.org/10.1371/journal.pone.0027272

Keysers (Social Brain Press, 2011). The Empathic Brain.

Etzel, Valchev & Keysers (2011) The impact of certain methodological choices on multivariate analysis of fMRI data with support vector machines. Neuroimage 54, 1159–67. https://doi.org/10.1016/j.neuroimage.2010.08.050

Bastiaansen, Meffert, Hein, Huizinga, Ketelaars, Pijnenborg, Bartels, Minderaa, Keysers & de Bildt (2011) Diagnosing autism spectrum disorders in adults: the use of Autism Diagnostic Observation Schedule (ADOS) module 4. J. Autism Dev. Disord. 41, 1256–66. https://doi.org/10.1007/s10803-010-1157-x

Keysers & Mckay (2011) How to make social neuroscience social. Psychol. Inq. 22, https://doi.org/10.1080/1047840X.2011.567960

Schippers, Renken & Keysers (2011) The effect of intra- and inter-subject variability of hemodynamic responses on group level Granger causality analyses. Neuroimage 57, 22–36. https://doi.org/10.1016/j.neuroimage.2011.02.008

Hurlemann, Walter, Rehme, Kukolja, Santoro, Schmidt, Schnell, Musshoff, Keysers, Maier, Kendrick & Onur (2010) Human amygdala reactivity is diminished by the β-noradrenergic antagonist propranolol. Psychol. Med. 40, 1839–1848. https://doi.org/10.1017/S0033291709992376

Thioux & Keysers (2010) Empathy: shared circuits and their dysfunctions. Dialogues Clin. Neurosci. 12, 546–52.

Keysers, Kaas & Gazzola (2010) Erratum: Somatosensation in social perception (Nature Reviews Neuroscience (2010) 11 (417-428)). Nat. Rev. Neurosci. 11, https://doi.org/10.1038/nrn2919

Keysers, Kaas & Gazzola (2010) Somatosensation in social perception. Nat. Rev. Neurosci. 11, 417–28. https://doi.org/10.1038/nrn2833

Schippers, Roebroeck, Renken, Nanetti & Keysers (2010) Mapping the information flow from one brain to another during gestural communication. Proc. Natl. Acad. Sci. U. S. A. 107, 9388–93. https://doi.org/10.1073/pnas.1001791107

Keysers & Gazzola (2010) Social neuroscience: mirror neurons recorded in humans. Curr. Biol. 20, R353-4. https://doi.org/10.1016/j.cub.2010.03.013

Kokal & Keysers (2010) Granger causality mapping during joint actions reveals evidence for forward models that could overcome sensory-motor delays. PLoS One 5, e13507. https://doi.org/10.1371/journal.pone.0013507

Nanetti, Cerliani, Gazzola, Renken & Keysers (2009) Group analyses of connectivity-based cortical parcellation using repeated k-means clustering. Neuroimage 47, 1666–77. https://doi.org/10.1016/j.neuroimage.2009.06.014

Bastiaansen, Thioux & Keysers (2009) Evidence for mirror systems in emotions. Philos. Trans. R. Soc. Lond. B. Biol. Sci. 364, 2391–404. https://doi.org/10.1098/rstb.2009.0058

Keysers (2009) Mirror neurons. Curr. Biol. 19, R971-3. https://doi.org/10.1016/j.cub.2009.08.026

Onur, Walter, Schlaepfer, Rehme, Schmidt, Keysers, Maier & Hurlemann (2009) Noradrenergic enhancement of amygdala responses to fear. Soc. Cogn. Affect. Neurosci. 4, 119–26. https://doi.org/10.1093/scan/nsn049

Keysers & Gazzola (2009) Expanding the mirror: vicarious activity for actions, emotions, and sensations. Curr. Opin. Neurobiol. 19, 666–71. https://doi.org/10.1016/j.conb.2009.10.006

Kokal, Gazzola & Keysers (2009) Acting together in and beyond the mirror neuron system. Neuroimage 47, 2046–56. https://doi.org/10.1016/j.neuroimage.2009.06.010

Perrett, Xiao, Barraclough, Keysers & Oram (2009) Seeing the future: Natural image sequences produce ‘anticipatory’ neuronal activity and bias perceptual report. Q. J. Exp. Psychol. (Hove). 62, 2081–104. https://doi.org/10.1080/17470210902959279

Gazzola & Keysers (2009) The Observation and Execution of Actions Share Motor and Somatosensory Voxels in all Tested Subjects: Single-Subject Analyses of Unsmoothed fMRI Data. Cereb. Cortex 19, 1239–1255. https://doi.org/10.1093/cercor/bhn181

Schippers, Gazzola, Goebel & Keysers (2009) Playing charades in the fMRI: are mirror and/or mentalizing areas involved in gestural communication? PLoS One 4, e6801. https://doi.org/10.1371/journal.pone.0006801

Etzel, Gazzola & Keysers (2009) An introduction to anatomical ROI-based fMRI classification analysis. Brain Res. 1282, 114–125. https://doi.org/10.1016/j.brainres.2009.05.090

Del Giudice, Manera & Keysers (2009) Programmed to learn? The ontogeny of mirror neurons. Dev. Sci. 12, 350–63. https://doi.org/10.1111/j.1467-7687.2008.00783.x

Jabbi & Keysers (2008) Inferior frontal gyrus activity triggers anterior insula response to emotional facial expressions. Emotion 8, 775–80. https://doi.org/10.1037/a0014194

Kukolja, Schläpfer, Keysers, Klingmüller, Maier, Fink & Hurlemann (2008) Modeling a negative response bias in the human amygdala by noradrenergic-glucocorticoid interactions. J. Neurosci. 28, 12868–76. https://doi.org/10.1523/JNEUROSCI.3592-08.2008

Keysers & Fadiga (2008) The mirror neuron system: New frontiers. Soc. Neurosci. 3, 193–198. https://doi.org/10.1080/17470910802408513

Thioux, Gazzola & Keysers (2008) Action understanding: how, what and why. Curr. Biol. 18, R431-4. https://doi.org/10.1016/j.cub.2008.03.018

Jabbi, Bastiaansen & Keysers (2008) A common anterior insula representation of disgust observation, experience and imagination shows divergent functional connectivity pathways. PLoS One 3, e2939. https://doi.org/10.1371/journal.pone.0002939

Etzel, Gazzola & Keysers (2008) Testing Simulation Theory with Cross-Modal Multivariate Classification of fMRI Data. PLoS One 3, e3690. https://doi.org/10.1371/journal.pone.0003690

Keysers & Gazzola (2007) Integrating simulation and theory of mind: from self to social cognition. Trends Cogn. Sci. 11, 194–6. https://doi.org/10.1016/j.tics.2007.02.002

Gazzola, van der Worp, Mulder, Wicker, Rizzolatti & Keysers (2007) Aplasics born without hands mirror the goal of hand actions with their feet. Curr. Biol. 17, 1235–40. https://doi.org/10.1016/j.cub.2007.06.045

Gazzola, Rizzolatti, Wicker & Keysers (2007) The anthropomorphic brain: the mirror neuron system responds to human and robotic actions. Neuroimage 35, 1674–84. https://doi.org/10.1016/j.neuroimage.2007.02.003

van der Gaag, Minderaa & Keysers (2007) Facial expressions: what the mirror neuron system can and cannot tell us. Soc. Neurosci. 2, 179–222. https://doi.org/10.1080/17470910701376878

Jabbi, Swart & Keysers (2007) Empathy for positive and negative emotions in the gustatory cortex. Neuroimage 34, 1744–53. https://doi.org/10.1016/j.neuroimage.2006.10.032

van der Gaag, Minderaa & Keysers (2007) The BOLD signal in the amygdala does not differentiate between dynamic facial expressions. Soc Cogn Affect Neurosci 2, 93–103. https://doi.org/10.1093/scan/nsm002

Keysers & Gazzola (2006) Towards a unifying neural theory of social cognition. Prog. Brain Res. 156, 379–401. https://doi.org/10.1016/S0079-6123(06)56021-2

Gazzola, Aziz-Zadeh & Keysers (2006) Empathy and the somatotopic auditory mirror system in humans. Curr. Biol. 16, 1824–9. https://doi.org/10.1016/j.cub.2006.07.072

Keysers, Xiao, Foldiak & Perrett (2005) Out of sight but not out of mind: the neurophysiology of iconic memory in the superior temporal sulcus. Cogn. Neuropsychol. 22, 316–32. https://doi.org/10.1080/02643290442000103

Keysers & Perrett (2004) Demystifying social cognition: a Hebbian perspective. Trends Cogn. Sci. 8, 501–7. https://doi.org/10.1016/j.tics.2004.09.005

Foldiak, Xiao, Keysers, Edwards & Perrett (2004) Rapid serial visual presentation for the determination of neural selectivity in area STSa. Prog Brain Res 144, 107–116.

Gallese, Keysers & Rizzolatti (2004) A unifying view of the basis of social cognition. Trends Cogn. Sci. 8, 396–403. https://doi.org/10.1016/j.tics.2004.07.002

Keysers, Wicker, Gazzola, Anton, Fogassi & Gallese (2004) A touching sight: SII/PV activation during the observation and experience of touch. Neuron 42, 335–46.

Wicker, Keysers, Plailly, Royet, Gallese & Rizzolatti (2003) Both of us disgusted in My insula: the common neural basis of seeing and feeling disgust. Neuron 40, 655–64.

Keysers, Kohler, Umiltà, Nanetti, Fogassi & Gallese (2003) Audiovisual mirror neurons and action recognition. Exp. brain Res. 153, 628–36. https://doi.org/10.1007/s00221-003-1603-5

Edwards, Xiao, Keysers, Földiák & Perrett (2003) Color sensitivity of cells responsive to complex stimuli in the temporal cortex. J. Neurophysiol. 90, 1245–56. https://doi.org/10.1152/jn.00524.2002

Keysers & Perrett (2002) What competition? Trends Cogn. Sci. 6, 119.

Keysers & Perrett (2002) Visual masking and RSVP reveal neural competition. Trends Cogn. Sci. 6, 120–125.

Kohler, Keysers, Umiltà, Fogassi, Gallese & Rizzolatti (2002) Hearing sounds, understanding actions: action representation in mirror neurons. Science 297, 846–8. https://doi.org/10.1126/science.1070311

Umiltà, Kohler, Gallese, Fogassi, Fadiga, Keysers, Rizzolatti, Umilta, Kohler, Gallese, Fogassi, Fadiga, Keysers & Rizzolatti (2001) I know what you are doing. a neurophysiological study. Neuron 31, 155–165. https://doi.org/S0896-6273(01)00337-3 pii

Gallese & Keysers (2001) Mirror neurons: A sensorimotor representation system. Behav. Brain Sci. 24,

Baker, Keysers, Jellema, Wicker & Perrett (2001) Neuronal representation of disappearing and hidden objects in temporal cortex of the macaque. Exp. brain Res. 140, 375–81.

Keysers, Xiao, Földiák & Perrett (2001) The speed of sight. J. Cogn. Neurosci. 13, 90–101.

Keysers, Diekamp & Güntürkün (2000) Evidence for physiological asymmetries in the intertectal connections of the pigeon (Columba livia) and their potential role in brain lateralisation. Brain Res. 852, 406–13.

Schall, Keysers & Kast (1999) Pharmacology of sensory gating in the ascending auditory system of the pigeon (Columba livia). Psychopharmacology (Berl). 145, 273–82.